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Head of Design - Developer Tools & AI Infrastructure

Job in San Francisco, San Francisco County, California, 94199, USA
Listing for: DesignX
Full Time position
Listed on 2026-10-09
Job specializations:
  • Software Development
    Software Architect, Full Stack Developer, Backend Developer, AI Engineer (Applied/Software)
Salary/Wage Range or Industry Benchmark: 180000 - 300000 USD Yearly USD 180000.00 300000.00 YEAR
Job Description & How to Apply Below

Greylock has a long history of partnering with category-defining enterprise companies, and we're excited to be working with an early-stage startup building a critical developer platform for the AI era.

As AI changes how software is built and operated, developers need an entirely new set of tools to understand, manage, and improve increasingly complex systems. This company is tackling one of the most important problems in that emerging stack, with an experienced founding team and strong early momentum. Much like the best developer platforms from Datadog to Stripe, the product experience itself has the potential to become a meaningful differentiator.

Powerful infrastructure needs to feel intuitive, fast, and remarkably easy to use.

The Role

We're looking for a Head of Design to own and define the product experience from the ground up. This is a highly hands‑on leadership role. You'll set the design vision and standards for the company, but you'll also be the person doing the work, designing core product experiences, prototyping new ideas, working through complex workflows, and partnering directly with the founders and engineering team to ship.

You'll have the opportunity to shape the full developer experience, from the core product UI to APIs, SDKs, and other surfaces developers interact with. You'll also establish the design language, systems, and principles that can scale with the company over time. This is an opportunity for someone who wants the scope and ownership of a design leader while remaining deeply involved in the craft.

* Please note, this role is 5 days a week in office, in San Francisco

You likely have:
  • Experience leading design for 0→1 B2B, infrastructure, developer, or technically complex products
  • Exceptional product thinking and an ability to turn complex technical concepts into intuitive experiences
  • Strong interaction and visual design craft
  • A track record of personally designing and shipping products — and a desire to remain hands‑on
  • The ability to move from user insight and product strategy through flows, prototypes, high‑fidelity design, and launch
  • Experience partnering closely with founders, product, and engineering
  • Strong judgment and the ability to establish a point of view on what great design should look like as the company scales

Experience with modern developer tools is particularly compelling. We'd love to see what you've built — whether that's production code, prototypes, side projects, Git Hub contributions, or experiments with new AI tools. Bonus points if you're already experimenting with LLMs, prompting, AI‑assisted prototyping, or new ways of designing and building software.

Further details about the company, founders, and product will be shared with qualified candidates.

About Greylock:

Greylock is an early‑stage investor in hundreds of remarkable companies, including Airbnb, Linked In, Dropbox, Workday, Cloudera, Facebook, Instagram, Roblox, Coinbase, and Palo Alto Networks, among others. More can be found about us here:

About the Greylock Recruiting Team:

As full‑time, salaried employees of Greylock, we provide free candidate referrals/introductions to active and upcoming investments to help them grow/succeed (as one of the many services we provide). Our recruiting team, combined, has over 125 years of in‑house recruiting experience at successful startups through FAANG’s and over 30 years of VC Talent.

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