Principal Engineer, AI Security

Your Impact at LILA

As a Principal Security Engineer focused on AI Security, you will define and drive the technical strategy for securing how AI is used across Lila's enterprise. You will operate as a senior individual contributor, partnering with IT and business teams to ensure safe and compliant adoption of AI tools and platforms.

While Lila builds AI-powered systems, this role is primarily focused on securing the use of third-party and internally deployed AI tools across the enterprise - ensuring sensitive data, intellectual property, and scientific workflows are protected as AI becomes deeply embedded in how work gets done.

What You'll Be Building

• Enterprise AI Security Strategy - Define and implement security controls and guardrails for the use of AI tools (e.g., LLM APIs, SaaS AI platforms, and internal AI services) across the organization.
• AI Gateway & Agentic Gateway Security - Design and implement AI gateway controls to manage and monitor access to external and internal AI systems. Secure agentic workflows by enforcing identity, authorization, tool-use constraints, and policy controls for autonomous or semi-autonomous agents.
• AI Red Teaming & Adversarial Testing - Conduct red teaming and adversarial testing focused on enterprise AI usage, including prompt injection, data exfiltration, jailbreaks, and abuse of connected tools and plugins.
• Data Protection for AI Usage - Develop and enforce controls to prevent sensitive data leakage through AI systems, including input/output filtering, data classification, tokenization, and secure handling of prompts, embeddings, and outputs.
• Multi-Layer AI Security (Network, Endpoint, Data) - Integrate AI security into existing enterprise security layers: network visibility and control over AI service access, API traffic inspection, and zero trust enforcement; endpoint security for developer machines, research environments, browsers, and plugins; data layer controls ensuring proper handling of sensitive data when interacting with AI systems.
• AI Threat Modeling (Enterprise Context) - Develop threat models focused on enterprise AI usage, including risks such as data leakage, prompt injection, model misuse, supply chain risks from AI vendors, and unauthorized agent actions.
• Vendor & Platform Security - Assess and guide secure adoption of third-party AI vendors and platforms, including evaluating data handling practices, model behavior, and integration risks.
• Incident Response for AI Usage - Define and support response approaches for AI-related incidents, such as sensitive data exposure, policy violations, or misuse of AI tools.
• Cross-Functional Technical Leadership - Partner with Legal, Compliance, IT, and Engineering to align AI usage with regulatory requirements, data governance policies, and responsible AI practices.
• Security Enablement - Contribute to internal guidance and education on safe AI usage, including secure prompting, data handling, and appropriate use of AI tools.
• Security Tooling & Implementation - Evaluate and implement tooling for AI security, including AI gateways, DLP integrations, monitoring solutions, and policy enforcement mechanisms.

What You'll Need to Succeed

• 8+ years of experience in information security, with strong expertise in enterprise, cloud, or application security.
• Hands-on experience designing and implementing security controls in enterprise environments.
• Familiarity with AI/ML systems and how modern AI tools (LLMs, copilots, APIs) are used in practice.
• Experience with cloud platforms (AWS/GCP), SaaS security, and zero trust architectures.
• Experience with data protection technologies (e.g., DLP, data classification, access controls).
• Practical experience with threat modeling, red teaming, or adversarial testing.
• Strong communication and influence skills across technical and non-technical stakeholders.

Bonus Points For

• Experience securing enterprise use of LLMs, copilots, or generative AI platforms.
• Familiarity with AI gateways, prompt filtering, or model interaction controls.
• Experience evaluating or securing third-party AI vendors and APIs.
• Background in regulated environments (biotech, healthcare, defense, or government).
• Experience with…